Chinese researchers identify a unique deep-sea tilefish, Branchiostegus sanae, linking it to the warrior princess of Studio Ghibli’s Princess Mononoke.
Warrior Princess Inspires Name of Newly Discovered Deep-Sea Fish

Warrior Princess Inspires Name of Newly Discovered Deep-Sea Fish
A new species of fish, named after a beloved anime character, showcases the intricate connection between culture and science.
In a fascinating blend of culture and science, a newly discovered deepwater fish has been named Branchiostegus sanae, in honor of San, the warrior princess from the acclaimed Studio Ghibli animated film Princess Mononoke. Chinese researchers identified this unique species, which is known for its distinctive facial markings resembling those of the film’s character, who has captivated audiences since its release in 1997.
Princess Mononoke, which grossed over 19 billion yen (approximately £100.4 million) in Japan, features San as a fierce protector of the forest, embodying themes of ecological balance and the struggle for harmony between nature and humanity. The researchers, from institutions including the South China Sea Institute of Oceanology, appropriately chose “sanae” for the fish's specific name as a tribute to San’s character.
Branchiostegus sanae is classified within the deepwater tilefish family, Branchiostegidae. These fish are typically found at impressive depths, with some species dwelling as far as 600 meters below the ocean's surface. The novelty of this discovery is underscored by the fact that only three new species in the Branchiostegus genus have been documented over the last 34 years, highlighting the significance of their find.
Lead author before the study, Haochen Huang, emphasized the film’s messages relating to human-nature interaction, stating, “We hope to echo this message through the naming.” As deepwater tilefish are often discovered in markets across East and Southeast Asia, this discovery not only enriches marine biodiversity but also provides unique context steeped in cultural heritage.
The specimens of the new fish species have been preserved in marine biological collections, positioning them for further scientific examination. This connection between profound storytelling and marine discovery showcases the increasingly interwoven fabric of art and science in today’s world.